All the roadbed plywood is 3/4". Most of it is cut about 8 - 10" wide for the roadbed. So it's either the plywood is changing dimensions or the rail. I'm inclined to think its the plywood. Last year, the basement reached the same temps, in the upper 60s, but I think the humidity might have been a bit higher. Its 26% down there now. The house is much dryer than it has ever been in the 3 years I've been living there.
Even if the vertical studs of the walls wanted to twist, I would think the panels of 3/8" plywood that is attached to them would bend somewhat and absorb the twist without putting strain on the roadbed that is laid across the tops of them.
First, an apology. I didn't look at your pictures closely enough. Let's try again. In one of your first posts, you said, 'new house'. I thought you meant that it was a house that had just been built. But here you say, 'three years', so it's not a brand new house, which means that the studs should have acclimatized. Even if this winter's humidity is lower, the studs shouldn't move much after three years, although they might move a bit, depending on what the wood is. But looking at the pictures of the warped track, it looks as though it's the peninsula that's the location of the problem. So I'll go back to my first diagnosis. The three peninsulas and the one wall that forms a triangle are new. What did you use for uprights? Especially the peninsulas, where you have uprights with what looks like two foot plywood brackets attached to them. Again, if the uprights twist even a bit it'll cause the brackets to follow, or try to. But the brackets can't twist, as they are held in place. So four things can happen. (1) The screws holding the brackets to the uprights can detach; (2) The railbed can detach; (3) The railbed can twist; (4) The railbed can buckle. I think that you're saying it's the last that's happened, and are attributing it to a change in the plywood's dimensions. I don't think that's the case. I think it much more likely that the dimensional timbers are unstable, rather than the plywood. Plywood is like a sheet of paper - you can't put a compound curve in it. But you CAN twist it, and I think that's what's happening here. It wouldn't take much twist to produce the effect you have - say 1/4" over a four foot length.
I'm pursuing this because I'm thinking of anyone building a layout like yours, or even much more modest, with the concomitant effort and expense. If I'm right in my diagnosis, those people can save themselves a world of worry by choosing their materials carefully. To put it bluntly, a lot of dimensional lumber these days is garbage. ('Dimensional' meaning a nominal 2" thick or greater.) It comes from young trees, it's unstable, and I wouldn't use it for anything but firewood. The greater the size the better it's likely to be, as you can't get a 20' 2x12 from a 50 year old tree, so large dimension lumber is apt to have fewer knots and be more stable. Finger jointed wood is better than solid, as is laminate. Fir is better than pine. If you MUST use solid 2x4, let it sit in your layout room for a year or more to acclimatize.